The smell may possibly be coming from your drains. Try putting baking soda down there and then pouring boiling hot war down there that should help eliminate the smell if that is where is coming from. Some people also recommend adding vinegar to that mix.
If you can smell mildew but cannot see it, you have to start looking for it in less obvious places. Perhaps on the underside of surfaces, perhaps behind fixtures or even in fans or vents. There may also be mildew behind drywall. That may take a moisture meter to find.